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ow that all ancient and modern dogs share a common ancestry and descended from an ancient, extinct wolf population that was distinct from any modern wolf lineage. Some studies have posited that all living wolves are more closely related to each other than to dogs, while others have suggested that dogs are more closely related to modern Eurasian wolves than to American wolves. The dog is a domestic animal that likely travelled a commensal pathway into domestication (i.e. humans initially neither benefitted nor were harmed by wild dogs eating refuse from their camps). The questions of when and where dogs were first domesticated remains uncertain. Genetic studies suggest a domestication process commencing over 25,000 years ago, in one or several wolf populations in either Europe, the high Arctic, or eastern Asia. In 2021, a literature review of the current evidence infers that the dog was domesticated in Siberia 23,000 years ago by ancient North Siberians, then later dispersed eastward into the Americas and westward across Eurasia, with dogs likely accompanying the first humans to inhabit the Americas. Some studies have suggested that the extinct Japanese wolf is closely related to the ancestor of domestic dogs. In 2018, a study identified 429 genes that differed between modern dogs and modern wolves. As the differences in these genes could also be found in ancient dog fossils, these were regarded as being the result of the initial domestication and not from recent breed formation. These genes are linked to neural crest and central nervous system development. These genes affect embryogenesis and can confer tameness, smaller jaws, floppy ears, and diminished craniofacial development, which distinguish domesticated dogs from wolves and are considered to reflect domestication syndrome. The study concluded that during early dog domestication, the initial selection was for behavior. This trait is influenced by those genes which act in the neural crest, which led to the phenotypes observed in mod